巕擷嬠偍傛傃晄姰慡巕擷嬠偺岎攝偲岎攝宆堚揱巕
Sexual and asexual development in ascomycetes and mating type genes
桳峕丂椡丒嬥巕丂岟*丒嬥怷惓庽**丒壛摗僴僫***丒堜嶳恀嬚****丒晉妦壛撧帥壀丂揙
Tsutomu Arie, Isao Kaneko*, Masaki Kanamori**, Hana Kato***, Makoto Iyama****, Kana Togashi and Tohru Teraoka

Abstract

巕擷嬠栧偵懏偡傞庬偵偼婎杮揑偵2庬椶偺岎攝宆乮MAT1-1偁傞偄偼MAT1-2偺偄偢傟偐乯偺嬠姅偑懚嵼偟丄堎側傞岎攝宆嬠姅偑岎攝偡傞偙偲偱桳惈惗怋傪峴偄丄姰慡悽戙傪宍惉偡傞乮self-incompatible = heterothallic乯丅奺嬠姅偺岎攝宆偼丄僎僲儉忋偺岎攝宆堚揱巕乮MAT1乯椞堟偵懚嵼偡傞懳棫堚揱巕孮乮MAT1-1堚揱巕孮偁傞偄偼MAT1-2堚揱巕孮乯偺偄偢傟傪帩偮偐偵傛偭偰寛掕偝傟傞丅 MAT1椞堟偺夝愅偼丄摉弶丄Neurospora crassa丄Podospora anserina丄Cochliobolus heterostropus乮僩僂儌儘僐僔偛傑梩屚昦嬠乯摍偱恑傔傜傟偨丅偄偢傟偺庬偵偍偄偰傕MAT1椞堟偵偼丄MAT1-1丄MAT1-2嬠姅娫偱墫婎攝楍偺憡摨惈偑擣傔傜傟側偄椞堟偑丄椉岎攝宆嬠姅偱嫟捠偺椞堟偵嫴傑傟偰懚嵼偡傞偙偲偑曬崘偝傟偨丅偙偺旕憡摨椞堟傪偦傟偧傟MAT1-1丄MAT1-2僀僨傿僆儌儖僼乮idiomorph乯偲屇傇丅僀僨傿僆儌儖僼忋偵偼丄偦傟偧傟丄DNA寢崌僪儊僀儞乮兛-box偍傛傃HMG-box乯傪帩偮MAT1-1-1偍傛傃MAT1-2-1堚揱巕偑懚嵼偡傞丅偦傟偧傟偺DNA寢崌僪儊僀儞偺傾儈僲巁攝楍偼庬傪挻偊偰曐懚惈偑崅偄偺偑摿挜偱偁傝丄偙傟傪棙梡偟偰懡庬偱MAT1椞堟偺夝愅偑恑傫偩丅堚揱巕攋夡幚尡摍偵傛偭偰丄MAT1-1-1偍傛傃MAT1-2-1偑僐乕僪偡傞僞儞僷僋幙偑丄嬠姅偺岎攝宆寛掕丄岎攝奐巒偺惂屼偵娭梌偡傞偙偲偑帵偝傟偰偄傞丅Co. hererostrophus偺MAT椞堟偵偼偙偺1慻偺堚揱巕偺傒偑丄N. crassa偍傛傃Po. anserina偱偼丄MAT1-1僀僨傿僆儌儖僼忋偵偝傜偵2偮偺堚揱巕MAT1-1-2丄MAT1-1-3偑懚嵼偡傞丅偙傟傜偺堚揱巕嶻暔偼柅惈摍丄桳惈惗怋偵娭梌偡傞偙偲偑帵嵈偝傟偰偄傞丅 偦偺屻丄Mycosphaerella graminicola丄Didymella rabiei丄Cryphonectria parasitica丄Gibberella fujikuroi丄Paecilomyces tenuipes丄Pyrenopeziza brassicae摍偺self-incompatible庬丄偍傛傃Co. homomorphus丄G. zeae摍偺self-compatible 乮= homothallic乯庬偱MAT1椞堟偺峔憿偑惛嵏偝傟偰偄傞丅徻偟偔偼Debuchy & Turgeon 2005傪嶲徠偄偨偩偒偨偄丅 巕擷嬠偼懡偔偺岎攝晄慡惈乮asexual乯偺庬傪娷傓丅姰慡悽戙宍惉偑枹曬崘偐偁傞偄偼杦偳尒傜傟側偄丄偄傢備傞晄姰慡巕擷嬠椶偱偁傞丅怉暔昦尨嬠傪娷傓Fusarium oxysporum丄Alternaria alternata丄Biporalis sacchari傗崓嬠Aspergillus oryzae偱偼姰慡悽戙偑枹曬崘偱偁傞丅摉弶丄偙傟傜偺庬偑岎攝晄慡偱偁傞尨場偑MAT1椞堟偺寚懝傗晄慡丄堦曽偺岎攝宆嬠姅偺寚擛偵偁傞偺偱偼側偄偐偲憐掕偝傟偨偑丄偦偺屻F. oxysporum丄A. alternata丄B. sacchari偵偍偄偰傕椉岎攝宆偺嬠姅偑懚嵼偡傞偙偲偑DNA儗儀儖偱妋擣偝傟丄偝傜偵丄偦傟傜偺MAT1椞堟偑嬤墢偺僿僥儘僞儕僢僋庬偲憡摨偱偁傞偽偐傝偐丄嵗忔偡傞MAT1-1-1偍傛傃MAT1-2-1堚揱巕偑敪尰偟丄婡擻惈傪傕曐帩偟偰偄傞偙偲偑帵偝傟偨丅偙偺偨傔丄岎攝晄慡偺尨場偼丄岎攝堚揱巕偺壓棳偱婡擻偡傞丄岎攝娭楢僔僌僫儖揱払宯偵偁傞偲憐掕偝傟偰偄傞丅 僀僱偺嵟廳梫昦奞偱偁傞偄傕偪昦偺昦尨嬠Magnaporthe grisea乮anamorph, Pyricularia grisea乯偼丄杮棃僿僥儘僞儕僢僋側岎攝傪峴偆偼偢偱偁傞偑丄拞崙塤撿側偳偱嵦廤偝傟偨堦晹偺姅傪彍偒丄捠忢丄岎攝擻傪帵偝側偄丅擔杮嶻偄傕偪昦嬠摨巑偺偐偗崌傢偣偵傛傞姰慡悽戙宍惉偺曬崘偼側偔丄捠忢丄岎攝晄慡惈偱偁傞偲棟夝偝傟偰偄傞丅偝偰丄偄傕偪昦嬠崙撪暘棧姅傪嫙帋偟夝愅傪峴偭偨寢壥丄変乆偼丄岎攝宆MAT1-1偍傛傃MAT1-2偺姅偑懚嵼偡傞偙偲丄夵傔偰MAT1-1嬠姅偲MAT1-2嬠姅傪偐偗崌傢偣偰傕姰慡悽戙傪宍惉偟側偄偙偲丄擔杮嶻姅偺MAT1椞堟偺峔憿偼僿僥儘僞儕僢僋側塤撿姅偲傎傏摨條偱偁傞偙偲丄MAT1椞堟忋偺堚揱巕乮MAT1-1-1丄MAT1-1-2丄MAT1-1-3偍傛傃MAT1-2-1丄MAT1-2-2乯偼擔杮嶻姅偱傕敪尰偟偰偄傞偙偲丄傪柧傜偐偵偟偨丅峔憿夝愅偺夁掱偱尒弌偟偨MAT1-1-3偍傛傃MAT1-2-1偼丄MAT1-1偍傛傃MAT1-2僀僨傿僆儌儖僼忋偐傜嫟捠攝楍偵偐偗偰懚嵼偡傞嫽枴怺偄峔憿傪偲偭偰偄偨丅偦偺偨傔丄椉堚揱巕偑僐乕僪偡傞嶻暔偼丄戝晹暘偺傾儈僲巁偑嫟捠偱丄N枛懁偺嬐偐側傾儈僲巁偑堎側傞偙偲偵傛偭偰丄岎攝宆摿堎惈傪帩偮丅偙偺嶻暔偺嵎偑婡擻揑偵偳偺傛偆側堄枴傪惗偠傞偺偐丄傑偨丄椉堚揱巕偑惗偠偨攚宨偵偮偄偰偼枹抦偱偁傞丅偝傜偵丄MAT1-1-3堚揱巕偺忋棳偵CT斀暅攝楍 (CT)n偑懚嵼偡傞偙偲丄偙偺CT偺斀暅悢偑嬠姅偵傛偭偰堎側傞乮僿僥儘僞儕僢僋姅偱戝偒偔丄岎攝晄慡姅偱彫偝偄乯偙偲丄CT斀暅悢偺懡偄姅偱MAT1-1-3偺揮幨検偑懡偄孹岦偵偁傞偙偲傪尒弌偟偨丅 岎攝宆堚揱巕嶻暔偼丄僼僃儘儌儞傗僼僃儘儌儞儗僙僾僞乕側偳偺嶻惗傪惂屼偟丄儗僙僾僞乕偑庴梕偟偨乽懠岎攝宆嬠姅偺懚嵼乿偲偄偆忣曬偑嶰検懱G僞儞僷僋幙丄MAP僉僫乕僛宱楬偁傞偄偼cAMP宱楬傪夘偟偰妀撪偵揱払偝傟丄嬠巺梈崌丄妀梈崌丄巕幚懱宍惉側偳偺岎攝嫇摦偑婲偙傞偙偲偑憐掕偝傟偰偄傞丅変乆偼丄僿僥儘僞儕僢僋庬偱偁傞Gibberella fujikuroi乮anamorph, Fusarium sacchari; 僒僩僂僉價偟傚偆摢晠攕昦嬠乯偵偍偄偰G僞儞僷僋幙兝僒僽儐僯僢僩偍傛傃MAP僉僫乕僛堚揱巕偺攋夡姅傪偦傟偧傟嶌弌偟丄偳偪傜偺攋夡傕嬠巺惉挿偵偼塭嬁傪媦傏偝側偄偙偲丄帗惈晄柅偲側傞偙偲丄昦尨惈偑掅壓偡傞偙偲傪尒弌偟偨丅崱屻偼偙傟傜傪娷傓忣曬揱払宯偺夝愅傪恑傔丄僿僥儘僞儕僢僋庬偲岎攝晄慡庬偺斾妑傪峴偆偙偲偱岎攝晄慡偺尨場傪扵傝偨偄偲峫偊偰偄傞丅
*尰 擾嬈娐嫬媄弍尋媶強
**尰 僒儞僩儕乕愭恑媄弍墳梡尋媶強
***尰TIS
****尰 JICA


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